Tayside Biodiversity - Tayside Biodiversity Action Plan - Water - Mesotrophic Lochs
INTRODUCTION
Mesotrophic lochs were identified by the UK Biodiversity Group as a key habitat of particular national importance that required specific work over and above that detailed in the standing open waters broad habitat plan. Much of the Standing Open Waters Habitat Action Plan therefore applies, but issues particularly relevant to mesotrophic lochs are examined here.

DEFINITION
Mesotrophic lochs are defined either as those with a moderately rich plant nutrient environment, particularly nitrogen and phosphorus, or those having a range of submerged plant communities, principally NVC (National Vegetation Classification) types 5A and 5B.

Type 5A is characterised by Shore-weed Littorella uniflora, Alternate flowered water milfoil Myriophyllum alterniflorum, Stoneworts Nitella spp., Small pondweed Potamogeton berchtoldii and Canadian pondweed Elodea canadensis, (an alien
species). Type 5B is characterised by Floating pondweed Potamogeton natans, and White water lilies Nymphaea alba.

However, lochs may be historically mesotrophic but have been subsequently changed to eutrophic by human activity. This type of loch has been included within this Plan as it may be possible in the long term to return them to a more natural nutrient status.
CURRENT STATUS AND EXTENT OF HABITAT
There are several mesotrophic lochs in Tayside, mainly located along the fringe of the uplands. These are listed under key sites, together with a brief assessment of their current status. It is apparent from the list that many lochs that were probably historically mesotrophic have now become eutrophic and others are threatened by nutrient enrichment. The Lowes chain of lochs between Dunkeld and Blairgowrie, which is of international significance, remains of high quality, but is threatened. Remediation of culturally eutrophic lochs is usually difficult, but efforts are being made, for example, to restore Loch Leven to a more natural nutrient status.

Naturally mesotrophic lochs are severely threatened and positive action is required to ensure the survival of this important habitat. Formerly mesotrophic lochs which have lost their characteristic animal and plant communities are not regarded as a national priority for remediation as it is considered that resources are best directed at maintaining and improving relatively unimpacted examples.

KEY SITES
The national rarity and decline of naturally mesotrophic lochs means that all of those present within the region merit inclusion as key sites. However, a few that particularly stand out are the Lowes chain of lochs between Dunkeld and Blairgowrie (Craiglush, Lowes, Butterstone, Clunie and Marlee), the Black Loch at Cleish, and Loch Moraig. These represent relatively unpolluted or pristine examples. Loch Leven, possibly once a mesotrophic loch, but now highly eutrophic, is of considerable importance and represents a classic example of the challenges faced in restoring lochs such as these to their natural status.
Loch Moraig (SSSI) -


Black Loch, Cleish (SSSI)


The Lowes chain of lochs of between Dunkeld and Blairgowrie of international
significance (SSSIs and increasing candidate SAC):


Loch of Lintrathen (SSSI)
Long Loch of Lundie (SSSI)
Crombie Reservoir
Loch Monzievaird
Drumore Loch (SSSI)
Laird’s Loch (SSSI)
Loch Leven (SSSI, SPA,
RAMSAR)
Rescobie Loch
Balgavies Loch
Mesotrophic, very high quality. A naturally mesotrophic loch, but there are indications that land use within the catchment is having an effect on the ecology of the loch.

Mesotrophic, very high quality. The only known naturally mesotrophic loch in Tayside that has had no recent agricultural improvement or afforestation within its catchment.

- Loch of Craiglush - oligo/mesotrophic, high quality.
- Loch of Lowes - mesotrophic, high quality, but indications of increasing nutrient levels.
- Butterstone Loch - mesotrophic, high quality, but indications of nutrient levels.
- Loch Clunie - mesotrophic, high quality, but indications of elevated nutrient levels.
- Loch of Drumellie or Marlee - mesotrophic, high quality, but indications of increasing nutrient levels.

Mesotrophic/eutrophic, artificial loch.
High quality, mesotrophic with notable fringing fen.
Artificial waterbody.
Status uncertain.
Mesotrophic loch.
Status uncertain.
Now eutrophic, may historically have been mesotrophic. Still of very high conservation value.
Eutrophic, may historically have been mesotrophic.
Eutrophic, may historically have been mesotrophic.
Slender Niaid

A Naiad, the water nymph and lake dweller of Greek myth, is found in Scotland...

An inconspicuous plant, the wonderfully named Slender Naiad Najas flexillis is typically found growing submerged in clear mesotrophic lochs where there is soft silty substrata.

Classed as ‘scarce’ in Britain, all the UK populations are now only found in Scotland. Most of these populations are found on the islands off the west coast, but there are a few sites on the mainland. Tayside’s only known population is found in the Lunan chain of lochs. It is thought to be in decline because of eutrophication, but it may also be due to excessive growth of other plants such as the invasive Canadian pondweed Elodea canadensis.
NATURE CONSERVATION IMPORTANCE
Mesotrophic lochs are frequently of particular nature conservation importance for the exceptionally diverse plant communities they support and for rare species such as Slender naiad Najas flexilis. Most mesotrophic lochs have SSSI status and the Lowes chain of lochs is a candidate Special Area of Conservation (SAC) for its populations of Slender naiad. These designations reflect the importance and scarcity of this habitat.

KEY SPECIES
Some species, such as the Slender naiad are very characteristic of mesotrophic lochs and their distribution is restricted
to these types of waterbodies. They, therefore, merit inclusion as key species for their nationally important populations.
Many of the key species associated with mesotrophic lochs such as the Common scoter Melanitta nigra, Black-necked
grebe Podiceps nigricollis and the Osprey Pandion haliaetus are also associated with other open standing water, but
mesotrophic lochs provide good habitat and may maintain significant populations.

P = UK Priority species C = UK species of conservation concern
Mammals Water vole Arvicola terrestris
P
Otter Lutra lutra
P
Daubenton’s bat Myotis daubentoni
C
Birds Common scoter Melanitta nigra
P
Osprey Pandion haliaetus
C
Black-necked grebe Podiceps nigricollis
C
Whooper swan Cygnus cygnus
C
Wigeon Anas penelope
C
Gadwall Anas strepera
C
Pink-footed goose Anser brachyrhynchos
C
Greylag goose Anser anser
C
Pochard Aythya ferina
C
Goosander Mergus merganser
C
Amphibians Palmate newt Triturus helveticus
C
Plants Slender naiad Najas flexilis
P
Slender stonewort Nitella gracilis
P
Pillwort Pilularia globulifera
P
NATIONAL BIODIVERSITY CONTEXT
There is a UK Habitat Action Plan for mesotrophic lochs. This has the following objectives:

 Maintain the characteristic plant and animal communities of current mesotrophic lochs.
 Identify and implement effective remedial action to address nutrient enrichment and pollution in mesotrophic lochs by 2010.

ECOLOGY AND MANAGEMENT

The ecology of mesotrophic lochs is critically dependent on nutrient levels. The diverse macrophyte communities they support are due to the excellent conditions they have for rooted plant growth. The moderate levels of nutrients they have encourage plant growth, whilst not generally being sufficient to cause algal blooms which can shut light out from rooted plants. Maintenance of natural nutrient levels is, therefore, a key element in the management of mesotrophic lochs. To achieve this a catchment based approach is required.
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