Tayside Biodiversity - Tayside Biodiversity Action Plan - Woodland - Upland Oakwoods
Wildcat (Felis sylvestris)
Ecology - Wildcats have increased in both numbers and range in Scotland in the past century and have benefited from afforestation. The species is found mainly in upper valley slopes and steep hillsides, on moorland and peatland and in rough grassland or forest, as well as in the lowland margins. They have also been recorded at higher altitudes, above 800m. It is listed as a ‘Species of Conservation Concern’ in Tayside.


Distribution - Widespread, except in the lowlands in the far east of Tayside. More abundant in Angus and east Perthshire. In recent decades there has been little change in overall range and the population in most areas appears stable.
There is a strong understorey and structure associated with this woodland holding a strong climax woodland community. Oakwoods and their associated open areas provide a rich and diverse habitat for a variety of invertebrates, including the Pearl-bordered fritillary Boloria euphrosyne. A large proportion of species listed in the following table are associated with the open ground areas of oakwoods.

KEY SPECIES
P = UK Priority species C = UK species of conservation concern

Mammals Red squirrel Sciurus vulgaris
P
Wildcat Felis sylvestris
C
Badger Meles meles
C
Brown long-eared bat Plecotus auritus
C
Pipistrelle bat Pipstrellus pipstrellus
C
Birds Wood warbler Phylloscopus sibilatrix
C
Spotted flycatcher Muscicapa striata
P
Redstart Phoenicurus phoenicurus
C
Great spotted woodpecker Dendrocopus major
C
Amphibian Great crested newt Triturus cristatus
P
Invertebrates Pearl-bordered fritillary Boloria euphrosyne
P
Hairy wood ant (Northern) Formica lugubris
P
Higher Plants Wild hyacinth Hyacinthoides non-scripta
C
Small cow-wheat Melampyrum sylvaticum
P
Wilson’s Filmy Fern Hymenophyllum wilsonii
C
NATIONAL BIODIVERSITY CONTEXT
In the UK, upland semi-natural oakwoods have declined by about 30 - 40% in area over the last sixty years as a result of replanting, mainly with introduced conifers, clearance for quarries or other developments and conversion to rough grazings. Recent changes have greatly reduced the amount of inappropriate management of these woodlands.

The objectives of the UK Biodiversity Action Plan for Upland Oakwood are:

 Maintain the existing area of upland oakwood (70,000 - 100,000 ha.) and improve its condition by a mixture of management for timber (predominately as low intensity high forest), as sheltered grazing and minimum intervention.
 Avoiding other habitats of high nature conservation value, expand the area of upland oakwood by about 10% on to currently open ground by some planting, but particularly by natural regeneration by 2005.
 Identify and encourage the restoration of a similar area of former upland oakwood that has been degraded by planting with conifers or invasion by Rhododendron Rhododendron ponticum.
In the context of Scotland, the estimated area of all upland oakwoods is in the region of 35,000 ha. of which ancient semi-natural woodland (ASNW) accounts for 24,000 ha. The Scottish targets are to initiate action in 80% of all HAP sites (i.e. 28,000 ha.) by 2004 and achieve favourable condition in over 50% by 2010. Within these figures the target is to initiate action in 100% of HAP sites occurring within SSSIs and SACs by 2004 and achieve favourable condition in over 70% by 2010.

The restoration figures for upland oakwood in Scotland are 3,000 ha. by 2010. This includes 10 ha. in Angus and 350 ha. in Perth & Kinross.

The expansion targets for upland oakwood in Scotland are 3,000 ha. by 2005. This includes 110 ha. in Angus and 280 ha. in Perth & Kinross.

ECOLOGY AND MANAGEMENT
Sessile oak Quercus petrea and Downy birch Betula pubescens dominate this woodland type. However where Quercus robur occurs, especially in eastern Scotland, it is usually of planted origin. The lack of oak dominance in woodland on higher slopes can often be attributed to past management and is not a true reflection of the natural species composition of this woodland type. However with increased altitude Birch becomes the dominant species. The field layer is dominated by three types namely grasses, bracken and ericoid sub-shrubs. The relative proportions of these vary with
soil type, shade, grazing and topography.

The oak woodlands are in general a mosaic of different NVC communities with W11 and W17 the major community types.

NVC W11 This is the most common oceanic type of woodland community with base-poor brown earth soils dominated by Oak species and Downy birch Betula pubescens. Where oak dominates, it forms a high forest with a closed canopy of tall oak with poor crown development derived principally from coppice and planted oak of unknown origin. Where birch is more dominant the canopy is more open. The occasional Ash Fraxinus excelsior may be found with, locally in Tayside, Rowan Sorbus aucuparia and Hazel Corylus avellana more common.

NVC W17 These communities are less frequent than the W11, but these woodlands have very acid shallow soils with Oak and Downy birch still the dominant species. Where the oak is dominant the trees are of a poorer quality and smaller than W11 oakwoods. Where the Birch is more dominant, Rowan is present as a co-dominant species. The commonest other species are Holly Ilex aquifolium though generally quite scarce, with Ash and Hazel which tend to be found in pockets of base-rich wet flushes.

The cessation of coppicing and the increased grazing pressure from both permitted grazing of domestic animals and browsing by wild animals has contributed to a decrease in the structural diversity of this woodland type. It must be pointed out that oakwoods that are not, or have never been, coppiced are not necessarily biodiversity poor. The removal of substantial amounts of timber throughout the early 20th century also contributed significantly to the changing composition and structure seen in Tayside’s upland oakwoods today.
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